Specialty/Research: Biochemistry, Molecular Genetics, Plant Physiology, Molecular Cell Biology, Phycology

Specialty／Research: Aquatic Ecotoxicology, Fish Ecology and Biology, Marine Ecology

Specialty／Research: Photosynthesis, Phycology, Plant Biochemistry and Physiology

Specialty／Research: Marine molluscus biology, Marine benthic invertebrates and ecology

Specialty／Research: Systematics, Phylogeny, Population Genetics and Ecology

Specialty／Research: Marine Biology, Molecular Evolution, Molecular Ecology

Specialty／Research: Population Genetics, Phylogeography

Specialty／Research: Phycology, Aquatic Ecology, Aquaculture

Specialty／Research: Biochemistry, Physiology, Coral Biology

Specialty／Research: Microbiology

Specialty／Research: Marine ecology, Sponges

Specialty/Research: Marine biology,Coral ecology,Climate change adaptation
Science education

Specialty／Research: Genomics, Transcriptomics, Evolution, Phylogenetics

Specialty／Research: Ecological engineering, Biogeochemistry

Specialty／Research: Quantitative Biology, Marine Ecology, Fisheries Stock Assessment, Fish Sclerochronology

Specialty／Research: Marine Invertebrates, Coral Systematics, Evolutionary Biology, Geomicrobiology

Specialty／Research: Comparative Biomechanics, Marine Ecology, Evolution, Bioinformatics